1. The importance of olfactory communication (using odors to communicate) in the way of life of deer[主语] was documented[谓语] by a study of captive adult male deer[方式状语] a few decades ago[时间状语], which noted that males rubbed their foreheads on branches and twigs, especially as autumn approached[定语从句].
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2. Scrapes consist of a clearing (about 0.5 meter in diameter) and shallow depression made by pushing aside the leaves covering the ground;[并列主句1] after making the scrape[时间状语], the deer typically urinates in the depression[并列主句2].
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